Patti Clifford Data Entry Person: Patti Clifford WRA Score 5

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1 Famil: Mrtaceae Taxo: Acca sellowiaa Som: Feijoa sellowiaa (O. Berg) O. Berg Orthostemo sellowiaus O. Berg (basiom) Commo Name: guavastee pieapple-guava Questioaire : curret Status: Assessor Approved Assessor: Patti Clifford Desigatio: EVALUATE Data Etr Perso: Patti Clifford WRA Score Is the species highl domesticated? =-3, =0 102 Has the species become aturalized where grow? 103 Does the species have weed races? 201 Species suited to tropical or subtropical climate(s) - If islad is primaril wet habitat, the (0-low; 1-itermediate; 2- High substitute "wet tropical" for "tropical or subtropical" high) (See Appedix 2) 202 Qualit of climate match data (0-low; 1-itermediate; 2- High high) (See Appedix 2) 203 Broad climate suitabilit (evirometal versatilit) 204 Native or aturalized i regios with tropical or subtropical climates 205 Does the species have a histor of repeated itroductios outside its atural rage? =-2,?=-1, =0 301 Naturalized beod ative rage = 1*multiplier (see Appedix 2), = questio Garde/ameit/disturbace weed =0, = 1*multiplier (see Appedix 2) 303 Agricultural/forestr/horticultural weed =0, = 2*multiplier (see Appedix 2) 304 Evirometal weed =0, = 2*multiplier (see Appedix 2) 305 Cogeeric weed =0, = 1*multiplier (see Appedix 2) 401 Produces spies, thors or burrs 402 Allelopathic 403 Parasitic 404 Upalatable to grazig aimals 405 Toxic to aimals 406 Host for recogized pests ad pathoges 407 Causes allergies or is otherwise toxic to humas 408 Creates a fire hazard i atural ecosstems 409 Is a shade tolerat plat at some stage of its life ccle 410 Tolerates a wide rage of soil coditios (or limestoe coditios if ot a volcaic islad) 411 Climbig or smotherig growth habit Prit Date: 8/23/2011 Acca sellowiaa (Mrtaceae) Page 1 of 6

2 412 Forms dese thickets 501 Aquatic =5, =0 502 Grass 503 Nitroge fixig wood plat 504 Geophte (herbaceous with udergroud storage orgas -- bulbs, corms, or tubers) 601 Evidece of substatial reproductive failure i ative habitat 602 Produces viable seed 603 Hbridizes aturall 604 Self-compatible or apomictic 605 Requires specialist polliators =-1, =0 606 Reproductio b vegetative fragmetatio 607 Miimum geerative time (ears) 1 ear = 1, 2 or 3 ears = 0, 3 4+ ears = Propagules likel to be dispersed uitetioall (plats growig i heavil trafficked areas) 702 Propagules dispersed itetioall b people 703 Propagules likel to disperse as a produce cotamiat 704 Propagules adapted to wid dispersal 705 Propagules water dispersed 706 Propagules bird dispersed 707 Propagules dispersed b other aimals (exterall) 708 Propagules survive passage through the gut 801 Prolific seed productio (>1000/m2) 802 Evidece that a persistet propagule bak is formed (>1 r) 803 Well cotrolled b herbicides =-1, =1 804 Tolerates, or beefits from, mutilatio, cultivatio, or fire 805 Effective atural eemies preset locall (e.g. itroduced biocotrol agets) =-1, =1 Desigatio: EVALUATE WRA Score 5 Prit Date: 8/23/2011 Acca sellowiaa (Mrtaceae) Page 2 of 6

3 Supportig Data: Morto, J.. Fruits of warm climates. J.F. [Is the species highl domesticated? No ]"Nowhere has the feijoa received more attetio tha i New Zealad. A Aucklad urserma itroduced 3 cultivars from Australia 'Coolidge', 'Choiceaa', ad 'Superba' about The remaied little kow util 1930 whe the feijoa was advertised as a orametal plat. Later, after improvemet b selectio ad amig of tpes with large, superior fruits ad their vegetative propagatio, small commercial platigs were made i citrus-growig areas of the North Islad." [o evidece of domesticatio that reduces ivasive abilit] WRA Specialist. Persoal Commuicatio. [Has the species become aturalized where grow?] NA WRA Specialist. Persoal Commuicatio. [Does the species have weed races?] NA USDA, ARS, Natioal Geetic Resources Program. Germplasm Resources Iformatio Network (GRIN) [Olie Database Idex]. Natioal Germplasm Resources Laborator, Beltsville, Marlad USDA, ARS, Natioal Geetic Resources Program. Germplasm Resources Iformatio Network (GRIN) [Olie Database Idex]. Natioal Germplasm Resources Laborator, Beltsville, Marlad Gilma, E.F./Watso, D.G.. Feijoa sellowiaa: Feijoa. Uiversit of Florida IFAS Extesio, Tuiso, J.T./Zimmer, N.G.. Success i cotrollig local alie plats i Hawaii Volcaoes Natioal Park i: Alie plat ivasios i ative ecosstems of Hawaii: maagemet ad research. Uiversit, Hoolulu USDA, ARS, Natioal Geetic Resources Program. Germplasm Resources Iformatio Network (GRIN) [Olie Database Idex]. Natioal Germplasm Resources Laborator, Beltsville, Marlad Morto, J.. Fruits of warm climates. J.F Tuiso, J.T./Zimmer, N.G.. Success i cotrollig local alie plats i Hawaii Volcaoes Natioal Park i: Alie plat ivasios i ative ecosstems of Hawaii: maagemet ad research. Uiversit, Hoolulu Radall, R.P.. Global Compedium of Weeds - Idex [Olie Database] Radall, R.P.. Global Compedium of Weeds - Idex [Olie Database] Radall, R.P.. Global Compedium of Weeds - Idex [Olie Database] Radall, R.P.. Global Compedium of Weeds - Idex [Olie Database]. [Species suited to tropical or subtropical climate(s) - If islad is primaril wet habitat, the substitute "wet tropical" for "tropical or subtropical"? High] Native rage: Brazil - Mias Gerais [s.], Paraa, Rio Grade do Norte, Rio Grade do Sul, Sata Cataria, Sao Paulo; Argetia - Misioes; Urugua. [Qualit of climate match data? High] Native rage: Brazil - Mias Gerais [s.], Paraa, Rio Grade do Norte, Rio Grade do Sul, Sata Cataria, Sao Paulo; Argetia - Misioes; Urugua. [Broad climate suitabilit (evirometal versatilit)? No] USDA Hardiess zoes: 8A-11. [Native or aturalized i regios with tropical or subtropical climates? Yes] Acca selowiaa was oe of the 41 exotic plats that was cotrolled i Hawaii Volcaoes Natioal Park, because it was spreadig. [Native or aturalized i regios with tropical or subtropical climates? Yes] Native rage: Brazil - Mias Gerais [s.], Paraa, Rio Grade do Norte, Rio Grade do Sul, Sata Cataria, Sao Paulo; Argetia - Misioes; Urugua. [Does the species have a histor of repeated itroductios outside its atural rage? Yes] Acca sellowiaa was first grow i Europe i the late 1800's. It was itroduced to the Riviera, Ital ad Spai i the late 1890's. I the earl 1900's plats were itroduced to Califoria ad Jamaica. Plats have bee itroduced to Chile, the Caribbea, New Zealad, Africa, Russia, Sicil, Portugal, Ital ad Eglad. [Naturalized beod ative rage? Yes] Acca sellowiaa was spreadig i Hawaii Volcaoes Natioal Park. The populatio was cotrolled with herbicide ad maual methods. [Garde/ameit/disturbace weed? No] No evidece. [Agricultural/forestr/horticultural weed? No] No evidece. [Evirometal weed? No] No evidece. [Cogeeric weed? No] No evidece. Prit Date: 8/23/2011 Acca sellowiaa (Mrtaceae) Page 3 of 6

4 Morto, J.. Fruits of warm climates. J.F WRA Specialist. Persoal Commuicatio. [Allelopathic?] Ukow. [Produces spies, thors or burrs? No] "The plat is a bush shrub 3 to 20 ft (0.9-6 m) or more i height with pale gra bark; the spreadig braches swolle at the odes ad white-hair whe oug. The evergree, opposite, short-petioled, blutl elliptical leaves are thick, leather, 1 1/8 to 2 1/2 i ( cm) log, 5/8 to 1 1/8 i ( cm) wide; smooth ad gloss o the upper surface, fiel vei ad silver-hair beeath." Morto, J.. Fruits of warm climates. J.F. [Parasitic? No] Mrtaceae WRA Specialist. Persoal Commuicatio. [Upalatable to grazig aimals? Ukow] Natioal Ceter for Biotecholog Iformatio. PubMed. U.S. Natioal Librar of Medicie, Bethesda, Marlad Specialized Iformatio Services, U.S. Natioal Librar of Medicie. TOXNET toxicolog data etwork [olie database]. Natioal Istitutes of Health, Morto, J.. Fruits of warm climates. J.F Messig R.. Maagig fruit flies o farms i Hawaii. Isect Pests. College of Tropical Agriculture ad Huma Resources Uiversit of Hawaii at Maoa, 4.pdf Natioal Ceter for Biotecholog Iformatio. PubMed. U.S. Natioal Librar of Medicie, Bethesda, Marlad Specialized Iformatio Services, U.S. Natioal Librar of Medicie. TOXNET toxicolog data etwork [olie database]. Natioal Istitutes of Health, [Toxic to aimals? No] No evidece of toxicit. [Toxic to aimals? No] No evidece of toxicit. [Host for recogized pests ad pathoges?]" Platig of feijoas has bee officiall discouraged i New South Wales ad Victoria, Australia, because the fruit is a prime host of the fruit fl." [Host for recogized pests ad pathoges?] Acca sellowiaa is a fruit fl host i Hawaii. "Fruit flies have become serious pests i Hawaii sice the first species was foud here i about The are widespread, occurrig from sea level to above 7000 ft elevatio, ad feed o hudreds of host plat species, ma of which are ecoomic crops." [No idicatio that Acca sellowiaa will substatiall impact curret cotrol efforts.] [Causes allergies or is otherwise toxic to humas? No] No evidece of toxicit. [Causes allergies or is otherwise toxic to humas? No] No evidece of toxicit WRA Specialist. Persoal Commuicatio. [Creates a fire hazard i atural ecosstems?] Ukow Morto, J.. Fruits of warm climates. J.F. [Is a shade tolerat plat at some stage of its life ccle?] Ca tolerate partial shade Plats for a Future Database. Acca sellowiaa. Plats for a Future Database, cca+sellowiaa [Is a shade tolerat plat at some stage of its life ccle?] Prefers light shade Morto, J.. Fruits of warm climates. J.F Gilma, E.F./Watso, D.G.. Feijoa sellowiaa: Feijoa. Uiversit of Florida IFAS Extesio, Morto, J.. Fruits of warm climates. J.F. [Tolerates a wide rage of soil coditios (or limestoe coditios if ot a volcaic islad)? Yes] "While the shrub is ofte said to be adapted to a wide rage of soil tpes ad i Eglad does well eve where there is a high chalk cotet, it actuall prefers rich orgaic soil ad is ot ver thrift o light or sad terrai. Some believe that a acid soil is best but the feijoa has doe well o soil with a ph of 6.2." [Tolerates a wide rage of soil coditios (or limestoe coditios if ot a volcaic islad)? Yes] Soil toleraces: sad; loam; slightl alkalie; acidic; well draied; well-draied. [Climbig or smotherig growth habit? No] Shrub WRA Specialist. Persoal Commuicatio. [Forms dese thickets?] Ukow. Prit Date: 8/23/2011 Acca sellowiaa (Mrtaceae) Page 4 of 6

5 Morto, J.. Fruits of warm climates. J.F Morto, J.. Fruits of warm climates. J.F Staples, G.W./Herbst, D.R.. A Tropical Garde Flora - Plats Cultivated i the Hawaiia Islads ad Other Tropical Places. Bishop Museum Press, Hoolulu, HI Morto, J.. Fruits of warm climates. J.F Morto, J.. Fruits of warm climates. J.F Morto, J.. Fruits of warm climates. J.F. [Aquatic? No] Shrub; terrestrial. [Grass? No] Mrtaceae. [Nitroge fixig wood plat? No] Mrtaceae. [Geophte (herbaceous with udergroud storage orgas -- bulbs, corms, or tubers)? No] Shrub; wood. [ Evidece of substatial reproductive failure i ative habitat? No] "The feijoa is ative to extreme souther Brazil, orther Argetia, wester Paragua ad Urugua where it is commo wild i the moutais." [Produces viable seed? Yes] "Geerall grow from seed." Plats for a Future Database. Acca sellowiaa. Plats for a Future Database, cca+sellowiaa [Produces viable seed? Yes] Propagate from seed WRA Specialist. Persoal Commuicatio. [Hbridizes aturall?] Ukow Morto, J.. Fruits of warm climates. J.F. [Self-compatible or apomictic?] Some cultivars are self-compatible Plats for a Future Database. Acca sellowiaa. Plats for a Future Database, cca+sellowiaa [Self-compatible or apomictic?] Plat is ot self-fertile Morto, J.. Fruits of warm climates. J.F Morto, J.. Fruits of warm climates. J.F Staples, G.W./Herbst, D.R.. A Tropical Garde Flora - Plats Cultivated i the Hawaiia Islads ad Other Tropical Places. Bishop Museum Press, Hoolulu, HI [Requires specialist polliators? No] Bees are the chief polliator. [Reproductio b vegetative fragmetatio? ] Propagatio is usuall b seed. Air laerig is also used. [Reproductio b vegetative fragmetatio? Propagate b seed, cuttigs or air laerig WRA Specialist. Persoal Commuicatio. [Reproductio b vegetative fragmetatio? ] Ukow. [uclear from the literature] Morto, J.. Fruits of warm climates. J.F. [Miimum geerative time (ears)? ] Plats produce fruit i 3-5 ears from seed WRA Specialist. Persoal Commuicatio. [Propagules likel to be dispersed uitetioall (plats growig i heavil trafficked areas)?] Ukow Morto, J.. Fruits of warm climates. J.F Staples, G.W./Herbst, D.R.. A Tropical Garde Flora - Plats Cultivated i the Hawaiia Islads ad Other Tropical Places. Bishop Museum Press, Hoolulu, HI [Propagules dispersed itetioall b people? Yes] Plat is grow for its fruit ad as a orametal. [Propagules dispersed itetioall b people? Yes] Widel dispersed i subtropical ad tropical regios as a orametal or fruit tree WRA Specialist. Persoal Commuicatio. [Propagules likel to disperse as a produce cotamiat? No] No evidece. Prit Date: 8/23/2011 Acca sellowiaa (Mrtaceae) Page 5 of 6

6 Gressler, E./Pizo, M.A./Morellato, P.C.. Poliizacao e dispersao de semetes em Mrtaceae do Brasil (Polliatio ad seed dispersal of Brazilia Mrtaceae). Revista Brasil Bota. 29: Gressler, E./Pizo, M.A./Morellato, P.C.. Poliizacao e dispersao de semetes em Mrtaceae do Brasil (Polliatio ad seed dispersal of Brazilia Mrtaceae). Revista Brasil Bota. 29: Gressler, E./Pizo, M.A./Morellato, P.C.. Poliizacao e dispersao de semetes em Mrtaceae do Brasil (Polliatio ad seed dispersal of Brazilia Mrtaceae). Revista Brasil Bota. 29: Gressler, E./Pizo, M.A./Morellato, P.C.. Poliizacao e dispersao de semetes em Mrtaceae do Brasil (Polliatio ad seed dispersal of Brazilia Mrtaceae). Revista Brasil Bota. 29: Gressler, E./Pizo, M.A./Morellato, P.C.. Poliizacao e dispersao de semetes em Mrtaceae do Brasil (Polliatio ad seed dispersal of Brazilia Mrtaceae). Revista Brasil Bota. 29: Morto, J.. Fruits of warm climates. J.F Morto, J.. Fruits of warm climates. J.F Tuiso, J.T./Zimmer, N.G.. Success i cotrollig local alie plats i Hawaii Volcaoes Natioal Park i: Alie plat ivasios i ative ecosstems of Hawaii: maagemet ad research. Uiversit, Hoolulu Morto, J.. Fruits of warm climates. J.F. [Propagules adapted to wid dispersal? No] Bird dispersed, flesh fruit. [Propagules water dispersed?] Ukow. [Propagules bird dispersed? Yes] Acca sellowiaa is dispersed b birds i the Brasilia forest. [Propagules dispersed b other aimals (exterall)? No] Acca sellowiaa is dispersed b birds. A flesh fruit. [o meas of exteral attachmet] [Propagules survive passage through the gut? Yes] Birds are dispersers of Acca sellowiaa i the Brazilia forest. [Prolific seed productio (>1000/m2)? Yes] "The 20-ear-old Riviera plat referred to above is said to have bore a crop of 2,000 fruits. There are usuall 20 to 40, occasioall as ma as 100, ver small, oblog seeds hardl oticeable whe the fruit is eate." [Evidece that a persistet propagule bak is formed (>1 r)? ]"The seeds will retai viabilit for a ear or more if kept dr." [Well cotrolled b herbicides? Yes] Acca sellowiaa was aturalizig i Volcaoes Natioal Park, Hawaii. A cut stump treatmet usig Tordo RTU was used o mature plats. A ear later o plats of this species were foud i the park. [Tolerates, or beefits from, mutilatio, cultivatio, or fire?] "The feijoa requires little care beod good soil preparatio before platig. Subsequet cultivatio is iadvisable because of the plat's shallow, fibrous root sstem which should be left udisturbed." WRA Specialist. Persoal Commuicatio. [Effective atural eemies preset locall (e.g. itroduced biocotrol agets)?] Ukow. Prit Date: 8/23/2011 Acca sellowiaa (Mrtaceae) Page 6 of 6