Common Name: VARIABLE-LEAF INDIAN-PLANTAIN. Scientific Name: Arnoglossum diversifolium (Torrey & Gray) H.E. Robinson. Other Commonly Used Names: none

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Common Name: VARIABLE-LEAF INDIAN-PLANTAIN Scientific Name: Arnoglossum diversifolium (Torrey & Gray) H.E. Robinson Other Commonly Used Names: none

Previously Used Scientific Names: Cacalia diversifolia Torrey & Gray; Mesadenia difformis Small; Mesadenia diversifolia (Torrey & A. Gray) Greene Family: Asteraceae/Compositae (aster) Rarity Ranks: G2/S2 State Legal Status: Threatened Federal Legal Status: none Federal Wetland Status: none Description: Perennial herb with stem to 6½ feet tall (200 cm), slightly grooved and angled. Lower leaves with blades up to 4 inches (10 cm) long, heart- or lance-shaped with rounded or squared-off bases, and shallowly lobed or coarsely toothed margins; leaf stalks 4-8 inches (10-20 cm) long. Upper stem leaves are smaller and more deeply toothed, with short or no leaf stalks; green on both surfaces. Flower heads about ½ inch (1.25 cm) high, held at the tips of small branches in a large, open cluster at the top of the stem; each head surrounded by small, greenish-white, winged bracts. Flowers 5 per head, white to lavender. Similar Species: Broadleaf Indian-plantain (Arnoglossum ovatum) has a round (not angled) stem, leaves whitened on the lower surface, and pale yellow flowers. Pale Indian-plantain (A. atriplicifolium) is an upland plant with a waxy stem, leaves whitened on the lower surface, and white flowers. Related Rare Species: Grooved-stem Indian-plantain (Arnoglossum sulcatum, Special Concern) occurs in sphagnum bogs and wet woodlands in Early, Decatur, and Lee counties. It resembles variable-leaved Indian-plantain but it has yellow flowers, and its lower leaves are oval, tapering gradually to the base and forming wings on the leaf stalk. Habitat: Openings in floodplain forests over limestone with clayey, basic soils and a canopy of hardwood trees and bald cypress. Life History: Little is known about pollination and seed dispersal in variable-leaf Indianplantain; it is likely pollinated by bees and other insects, and its seeds are probably dispersed by a combination of wind, gravity, and a variety of insects and small animals. Each fruit is surrounded by a whorl of fine bristles, the pappus, which aids in wind dispersal. Plants probably flower the second year following seed germination. Survey Recommendations: Surveys are best conducted during flowering (May August) and fruiting (July September). Range: Georgia, Florida, and Alabama.

Threats: Ditching, draining, and filling in wetlands. Clearcutting in bottomlands and floodplains. Impounding streams to form ponds and lakes. Georgia Conservation Status: Nine populations are known, none on conservation lands. Conservation and Management Recommendations: Avoid logging, ditching, and draining in wetlands. Avoiding stream impoundment. Selected References: Allison, J.R. 1991. The status of Cacalia diversifolia and Lythrum curtissii in southwest Georgia. Georgia Natural Heritage Program, Social Circle. Chafin, L.G. 2007. Field guide to the rare plants of Georgia. State Botanical Garden of Georgia and University of Georgia Press, Athens. Cronquist, A. 1980. Vascular flora of the southeastern United States, Vol. 1, Asteraceae. University of North Carolina Press, Chapel Hill. FNA. 2006. Flora of North America. Vol. 20, Magnoliophyta: Asteridae, Part 7: Asteraceae, Part 2. Oxford University Press, New York. Godfrey, R.K. and J.W. Wooten. 1981. Aquatic and wetland plants of southeastern United States, Vol. 2, dicotyledons. University of Georgia Press, Athens. Harper, R.M. 1905. Phytogeographical explorations in the coastal plain of Georgia in 1903. Bulletin of the Torrey Botanical Club 32(3): 141-171. Kral, R. 1983. A report on some rare, threatened, or endangered forest-related vascular plants of the South. Technical Publication R8-TP2. United States Forest Service, Atlanta. Kral, R. and R.K. Godfrey. 1958. Synopsis of the Florida species of Cacalia (Compositae). Quarterly Journal of the Florida Academy of Sciences 21: 193-206. NatureServe. 2007. NatureServe Explorer. Arlington, Virginia. http://www.natureserve.org/explorer Patrick, T.S., J.R. Allison, and G.A. Krakow. 1995. Protected plants of Georgia. Georgia Department of Natural Resources, Natural Heritage Program, Social Circle. Weakley, A.S. 2007. Flora of the Carolinas, Virginia, Georgia, and surrounding areas. University of North Carolina Herbarium, Chapel Hill. Author of species account: Linda G. Chafin Date Compiled or Updated: L. Chafin, Sept. 2007: original account K. Owers, Jan. 2010: updated status and ranks, added pictures

Inflorescence

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