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FM 9(4) :Field mycology 24/7/08 12:32 Page 126 MACROFUNGI OF THE KENWOOD ESTATE Andy Overall* As many readers of FM may be aware, following previous articles (FM 7(3):82-84; 8(1):24-27), Hampstead Heath is my home patch for fungi. Recording of the macro fungi of the Heath by Fungi to be with /London Fungus Group, reached its 10th year in 2007 with 433 records to date. At 791 acres ( 320 Hectares) straddling 2 boroughs, that of Camden & Barnet, Hampstead Heath is far from being an easy place to cover, therefore it is understandable that some areas tend to be more neglected than others, for whatever reason, be it distance or the likelihood of any interesting fungi turning up. During 2007, apart from my usual forays upon the Heath, I decided to pay a lot more attention to the Kenwood Estate, as much, but not all of this area, had been rather neglected by LFG forays and personal visits by myself. Kenwood makes up 112 acres (45 Hectares) of the total of Hampstead Heath and is quite different from the rest of the Heath. This whole area was in fact designed by the 18th century landscape garden designer, Humphry Repton, complete with lakes, woods, meadows, farm, dairy, stables and a kitchen garden, so it is no surprise that it takes on a different air to that of the heath itself. Today it is managed by English Heritage having taken over from the Corporation of London in 1989; this means that the area is likely to receive more attention than it would have done prior to 1989. Fig. 1. Kenwood House, the view from Beech Mount. Photograph Andy Overall. * Flat 2, 39 North End Road, Golders Green, London NW11 7RJ 126

FM 9(4) :Field mycology 24/7/08 12:32 Page 127 Sprawling at the rear of the Kenwood House itself is a large area of short, lush green grass, known as The Pasture Ground which was once grazed by cattle during the 18th century. In front and to the side of the house two large lawns look ripe for Waxcaps and other unimproved grass-loving species. The house is definitely worth a visit for its paintings by Turner, Rembrandt, Reynolds, Gainsborough and Vermeer as well the wonderful decor & rooms. There are 2 ancient woodland remnants on the estate, both with SSSI status, the South Wood that has good populations of both Pedunculate and Sessile oak along with some beech and birch and the smaller North Wood with its towering oaks & beech trees on Bagshot sand, there are also unimproved meadows & bogs with mature Alder. Both remnants are of a wood that existed before 1600 and management by man, therefore earning the tag of being ancient woodland. The SSSI status of both woods were bestowed not only because of the age of the woods but for the many hundreds of invertebrates that they support via the dead, standing and fallen wood. Maybe it was because of this less wild nature of Kenwood that it wasn t a priority for me when looking for fungi; if so, I couldn t have been more wrong and should have known better. I have mentioned that not all of the estate had been ignored, areas I had visited were mainly the woods, South & North and other wooded areas along the Western Boundary with the Spaniards Road, namely Mount Tyndal & The Elms. It was some of the more open areas where meadow meets tree line that were to be the most interesting. At the northern end, on the eastern flank of the estate, just outside the fenced boundary, an area known as the Stable Field, fringed with Oak & Beech was to reveal some real gems. Due to the hot weather during April followed by the incessant rains of May onward, species of Amanita & Russula quickly took advantage so that by mid May Fig. 2. Russula zvarae showing its striking matt pink pileus and stipe. A nationally rare species it was a new record for the estate. Photograph Andy Overall. 127

FM 9(4) :Field mycology 24/7/08 12:32 Page 128 Amanita rubescens, A. excelsa var. spissa and A. fulva, had all made an appearance, mainly in a small open, wooded, valley-like area known as the Pasture Ground Copse, of mainly oak and birch. A. spissa became particularly prevalent during June. It was during the summer months that some of the more interesting discoveries began to emerge. Along the western edge of the stable field, during mid June, good collections of Russula virescens and Russula graveolens were made, the former being a new record to the heath. R. virescens is a species I usually associate with beech, here it was with oak, which is not unknown. There were several variations of R. graveolens present, as colouration varied from the usual reddishbrown to a vibrant purple-violet-brown. Here also in healthy numbers were large, robust, green-brown fruitbodies of R. heterophylla. Not that any of these species are unusual, it was just good to see good healthy populations of each, together at the same time. The outstanding species from this area was undoubtedly the rare, small to mediumsized Russula zvarae (Fig. 2) which appeared during early August in a drier period, on totally worn, exposed, dry and cracked, bare soil, around 10 feet from a huge oak tree. Not a species I was at all familiar with, even though it had a pink, matt/velvety cap and pink flushed stem, it had me somewhat flummoxed for a short while, until I got it under the microscope where it revealed characteristics to help tell it apart from similar species such as R. rosea: encrusted fushinophile hyphae and flexuous, branched hairs in the pileipellis and subglobose spores with relatively low warts and an incomplete reticulum. Other Russula species in this area included R. pseudointegra, R. amoenolens, R. grisea, R. ionochlora & the ubiquitous R. parazurea. Among the boletes along this narrow stretch was a new record for the heath of Leccinum crocopidium appearing during July. Also present were Leccinum scabrum, Boletus radicans, Boletus edulis, Boletus luridiformis, Boletus badius, Boletus pulverulentus, Gyroporus castanea and Xerocomus rubellus, At the base of one of the older beech trees on the stable field, Podoscypha multizonata occurs (it is one of the sites covered in a previous article on this species), this year was no exception with it fruiting at the end of August, though looking rather stunted and malformed in its growth. At the base of one of the few Scots Pine occurring on the estate, the rather resplendent and alluring Phaeolus schweinitzii was in evidence, another new record for the heath. Just to the east of stable fields, heading toward Hampstead Lane, a rather ramshackle set of wooden gates lead you into an area called the Kitchen garden where herbs and various shrubs are grown, it is also home to a wonderfully fruitful Mulberry tree. It was here on the edge of the lawn that four tightly knit clusters of the infrequent Psathyrella multipedata (Fig. 3) glistened with the early morning dew, first appearing at the end of August and then again in October. Moving away from the stable fields area and out onto the pasture ground a large grassland area fringed with various trees, including English oak, Holm oak, beech, hornbeam, lime and birch. Throughout the summer various Russula species came and went, including R. atropurpurea (one specimen of which was practically white/cream in colour), R. farinipes (new record), R. puellaris (new record), R. exalbicans, R. aeruginea, R. pseudointegra & R. pectinata. The most exciting record on the pasture ground was that of Amanita crocea, a first for the heath, a lone specimen sitting right out in the open, among short turf, a few feet away from two lone birch trees never orange in my opinion (more peach?), with reference to the vernacular name for this species. A slightly more robust species than A. fulva and possibly more tawny in colour, with a white, often greying, thicker volval sac as opposed to the thin fleshed & russet toned volval sac of A. fulva. Also in association with the same birch trees, though later in September, 128

FM 9(4) :Field mycology 24/7/08 12:32 Page 129 Leccinum scabrum came through, as did Russula aeruginea. Sandwiched between the pasture ground and the south wood are two ponds, 1000 Pond and Wood Pond. On the east side of 1000 pond is the sham bridge, a fake bridge, behind which are a row of alder trees. On the banks of the pond directly beneath the alder & the bridge, fairly large groups of Naucoria striatula were fruiting during mid October. In the South Wood, at the base of Oak trees, Grifola frondosa came and went during the month of September at four separate sites. Laetiporus sulphureus was widespread across the estate, mainly during July & Aug, mostly on oak, though one fruiting occurred on hawthorn, a first for me. Fistulina hepatica was also widespread on oak, mainly during September. Heading west of the pasture ground, toward the Spaniard Road, is the West Meadow where there is a small boggy area, called The North Copse that has a small group of mature alder trees mixed with silver birch. I made a few visits to this spot throughout the summer, finding little except for the odd earth-ball. It was during late September that two notable species appeared among the wet moss and soil directly beneath the Alder: Lactarius obscuratus (Fig. 4) and Laccaria purpureobadia. With L. obscuratus being so small and far from forthcoming with any latex, I was looking elsewhere for a short while, but when the white milk eventually showed on some specimens (although not all), I knew then what I was actually dealing with. Both of these species are new records for the heath. Running parallel with the ridge of the North Wood, is the West Lodge Drive, during October, sitting tight up against chestnut paling on the Bagshot sand were small groups of the Almond Fibrecap - Inocybe hirtella var. bispora (Fig. 5). I found the smell of almonds barely evident on fresh material, it wasn t until collected specimens had sat for a few hours in a container that it became clear and fairly potent. Fresh specimens had a different, rather unpleasant smell. Fig. 3. Psathyrella multipedata is frequently found in grassy areas around trees. The large clumps arise from a common rooting base. Photograph Andy Overall. 129

FM 9(4) :Field mycology 24/7/08 12:32 Page 130 Fig. 4. Lactarius obscuratus, one of a suite of Lactarius species associated with alder. It has smaller spores and lacks the olivaceous tinges of the closely related and equally common L. cyathuliformis. Photograph Andy Overall. Fig. 5. Inocybe hirtella var. bispora, a species which smells strongly of almonds when enclosed for a while in a container. Photograph Andy Overall. 130

FM 9(4) :Field mycology 24/7/08 12:32 Page 131 Fig. 6. Ramaria stricta is one of the few common species in the UK, often appearing on woodchip mulch. Photograph Andy Overall. The lawns around the house proved a little disappointing, as I was hoping for a good display of waxcaps, only two species turned up, the common Hygrocybe virginea and the uncommon H. citrinovirens.there were also a few difficult species of Inocybe that escaped Identification. There was a nice surprise however when I found Suillus luteus fruiting with a 7 year old Bhutan pine. The lawn that turned out to be more interesting was not one of those surrounding the house but a lawn that goes with the tied cottages belonging to the estate aptly named Cottage Lawn, which is close to the West lodge car park. At the far end of the lawn is a large Weeping willow. In late October, beneath the willow and all around its fringe, were literally hundreds of fruit bodies of an Entoloma species. Having looked it at under the microscope a few times and using the Noordeloos key in FAN 1 my conclusion was E. myremecophilum. Also found on the lawn were Rickenella fibula and R. swartzii I was really surprised not to find any Hygrocybe here but I suspect there will be in future visits. To finish off, growing on wood-chips, around the shrubbery, close to the Mansion Cottage visitor centre, Ramaria stricta (Fig. 6) was fruiting generously, unseen by most visitors except for the ever searching eyes of the field mycologist. 150 species of macro fungi were recorded for Kenwood from April to Nov 2007, some of which were new records for both the county and the Heath, I shall be paying a lot more attention to this area in the future. 131