Leaf Key for Common Broadleaf Trees in Maryland

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Leaf Key for Common Broadleaf Trees in Maryland By: Madeline Koenig & Kerry Wixted January 2016

Tree Guide Key I. Leaves alternate A. Leaves simple 1. Margins entire......................................... Pg. 1 a. Leaves oval to elliptic.............................. Pg. 1 b. Leaves not oval or elliptic.......................... Pg. 2 i. Leaves lobed................................. Pg. 2 ii. Leaves not lobed............................. Pg. 5 2. Margins toothed....................................... Pg. 6 a. Leaf bases uneven................................. Pg. 6 b. Leaf bases even................................... Pg. 7 i. Leaves lobed................................. Pg. 7 ii. Leaves not lobed............................. Pg. 7 B. Leaves compound.......................................... Pg. 10 II. Leaves opposite A. Leaves simple 1. Margins entire...................................... Pg. 13 a. Leaves lobed...................................... Pg. 13 b. Leaves not lobed.................................. Pg. 13 2. Margins toothed...................................... Pg. 14 a. Leaves lobed...................................... Pg. 14 b. Leaves not lobed.................................. Pg. 14 B. Leaves compound.......................................... Pg. 15 i

How to Use this Guide This guide was created as an easy-to-use reference for beginner tree identification. It is not a comprehensive guide. Trees represented in the guide include those that are native to Maryland, those that are introduced (exotic), and those that are exotic and create ecological problems (invasive). To use this guide, begin with the key on the previous page. Terms used in the key are illustrated on pages x - xii. Please note that leaves can be highly variable in shape, size, appearance, and color. Leaves selected for this guide are intended to represent commonly found phenotypes. Leaves represented are not shown to scale. Under each leaf, the following information will be listed: Simple and compound refer to whether or not a leaf the leaf is divided. Simple leaves have a single blade attached to a stem known as a petiole. Compound leaves have multiple blades known as leaflets. To make sure you are looking at a leaf and not a leaflet, look for the lateral bud where the petiole meets the twig. All leaves have this trait, but leaflets do not. This is a lateral bud. Common Name (Scientific name), Exotic or Invasive Distinguishing characteristics Common Identification Terms: Simple Versus Compound SIMPLE COMPOUND ii

Alternate Versus Opposite Leaves Alternate and opposite refer to the arrangement of leaves along the twig. Do the leaves appear to zig-zag along the twig, or are they directly across from one another? ALTERNATE OPPOSITE Entire Versus Toothed Leaf Margins Entire and toothed refer to the edge of the leaf. The edge of a leaf is also known as a margin. Do the leaf edges seem to zig-zag, or are they smooth? ENTIRE TOOTHED iii

Even Versus Uneven Leaf Bases Even and uneven refer to the shape of the base of the leaf. Do both sides of the base of the leaf line up with one another; or are they uneven? EVEN LEAF BASE UNEVEN LEAF BASE Lobed Versus Not Lobed Leaves Lobed and not lobed refer to the shape of the leaf. Does the leaf have sections that jut out, or does it have a consistent leaf edge? This is a lobe. LOBED NOT LOBED iv

Alternate, Simple, Entire, Oval - Elliptic Southern Magnolia (Magnolia grandiflora) Exotic Underside orange and hairy Evergreen Umbrella Magnolia (Magnolia tripetala) Leaves 10-24 Common Pawpaw (Asimina triloba) Leaves 6-12 Edible fruit Sweetbay Magnolia (Magnolia virginiana) Underside white Common Persimmon (Diospyros virginiana) Bark broken in small squares Edible fruit Pg. 1

Alternate, Simple, Entire, Oval Elliptic; Cont d Bradford Pear (Pyrus calleryana) Invasive White flowers with unpleasant smell in spring Small, round fruit in fall Rhododendron sp. (Rhododendron sp.) /Exotic Variable leaf shape Evergreen Spicebush (Lindera benzoin) Shrub Aromatic when crushed Red berries in fall Sassafras (Sassafras albidum) Hairy leaves Variable leaf shape (see pg. 4) Aromatic when crushed Black Gum (Nyssa sylvatica) Dark, shiny leaves Underside lighter Blocky bark Pg. 2

Alternate, Simple, Entire, Not Oval, Lobed All leaves on this page have bristle-tips. Scarlet Oak (Quercus coccinea) Twigs reddish-brown Buds hairy, pointed Tufts of hair on underside Inner-bark orangish-pink Southern Red Oak (Quercus falcata) Top lobe elongated Leaf base rounded Blackjack Oak (Quercus marilandica) Leaf lobes broad Leaf is triangular Pin Oak (Quercus palustris) Deep lopes Twigs shiny Buds light brown, hairless Inner bark pink Northern Red Oak (Quercus rubra) Variable leaf shape Leaves never hairy Twigs reddish-brown Buds hairy, pointed Inner bark pink Black Oak (Quercus velutina) Hairy near veins on underside Twigs light brown Buds tan, oval-shaped, hairy Inner bark orange or yellow Pg. 3

Alternate, Simple, Entire, Not Oval, Lobed; Cont d All leaves on this page do not have bristle-tips. Chestnut Oak (Quercus montana) Shallow lobes Post Oak (Quercus stellata) Big lobes Leaf is cross-shaped. White Oak (Quercus alba) Deeply lobed Leaf shape variable Grayish-green on top Sassafras (Sassafras albidum) Hairy leaves Leaf shape variable (see pg. 2) Aromatic when crushed Tulip Poplar (Liriodendron tulipfera) Has 4 lobes Flowers resemble tulips Pg. 4

Alternate, Simple, Entire, Not Oval, Not Lobed Ginkgo (Ginkgo biloba) Exotic Fan-shaped Commonly planted Eastern Redbud (Cercis canadensis) Heart-shaped Flowers pink-purple, present before leaves Has pea-like pods in fall Willow Oak (Quercus phellos) Narrow with bristle-tip Pg. 5

Alternate, Simple, Toothed, Leaf Bases Uneven American Basswood (Tilia americana) Smooth to touch Finely toothed Small fruits with leafy wings in fall American Elm (Ulmus americana) Mostly smooth to touch; Slippery Elm sandpapery with hairy underside Tree is vase-shaped Commonly planted Siberian Elm (Ulmus pumila) Invasive Leaves narrow Leaves up to 3 in length Fruit round, winged Mulberry sp. (Morus sp.) /Invasive Leaf shape variable (see pg. 7,9) Fruit resembles blackberries Milky sap Witch-hazel (Hamamelis virginiana) Yellow flowers in fall Pg. 6

Alternate, Simple, Toothed, Leaf Bases Even, Lobed American Sycamore (Platanus occidentalis) Leaves large, broad Bark gray, white, pale green Sweetgum (Liquidambar styraciflua) Leaves star-shaped; Tree produces spiky, ballshaped seed pods in fall Mulberry sp. (Morus sp.) /Invasive Leaf shape variable (see pg. 6,9) Fruit resembles blackberries Milky sap Alternate, Simple, Toothed, Leaf Bases Even, Not Lobed River Birch (Betula nigra) Bark peeling, bright orange or reddish-brown Hophornbeam (Ostrya virginiana) Buds round Leaf veins sometimes forked Fruit round, winged Ironwood American Hornbeam (Carpinus caroliniana) Buds square Leaf veins never forked Fruit winged Bark smooth, gray Musclewood Pg. 7

Alternate, Simple, Toothed, Leaf Bases Even, Not Lobed; Cont d American Chestnut (Castanea dentata) Leaves long, narrow Leaf base tapers Feels thin, papery Chinese Chestnut (Castanea mollissima) Exotic Leaves oval Leaf base rounded Feels thick, waxy American Beech (Fagus grandifolia) Bark smooth, gray Buds slender, golden, pointed Black Cherry (Prunus serotina) Underside has hair along the main leaf vein Two small glands located near leaf base Japanese Flowering Crabapple (Malus floribunda) Exotic Flower small, white or pink in spring Fruit round, red or yellow in fall Pg. 8

Alternate, Simple, Toothed, Leaf Bases Even, Not Lobed Cont. Hawthorn sp. (Crataegus sp.) Flowers small, white with unpleasant smell in spring Fruit reddish in fall Wild varieties have thorns American Holly (Ilex opaca) Leaves dark, shiny, prickly Evergreen Persistent red berries in winter Mulberry sp. (Morus sp.) /Invasive Leaf shape variable (see pg. 7) Fruit resembles blackberries Milky sap Eastern Serviceberry (Amelanchier canadensis) White flowers in spring Underside lighter Crape Myrtle (Lagerstroemia indica) Exotic Bark peeling, smooth Flower color variable Willow sp. Salix sp. /Exotic Leaves narrow, elongated Pg. 9

Alternate, Compound Black Walnut (Juglans nigra) 7-23 leaflets Nuts round with green husk, woody shell Bark deeply furrowed Tree of Heaven (Ailanthus altissima) Invasive Leaflets mostly smooth, few teeth near base Seeds have papery wings Smooth Sumac (Rhus glabra) Leaflets toothed Twigs flat-sided, not hairy Reddish berries in upright cluster in winter Staghorn Sumac (Rhus typhina) Leaflets are toothed Twigs hairy Reddish berries in upright cluster in winter Winged Sumac (Rhus copallina) Leaflets not toothed Twigs hairy Leafstalks winged Reddish berries in upright cluster in winter Pg. 10

Alternate, Compound Continued Bitternut Hickory (Carya cordiformis) 7-11 leaflets Twigs somewhat hairy Buds yellow, powdery Mockernut Hickory (Carya tomentosa) 7-9 leaflets Twigs reddish brown, hairy End buds more than ½ long Nut husk thick Shagbark Hickory (Carya ovata) 5-7 leaflets Twigs reddish-brown, smooth End buds more than ½ long Bark shaggy Nut husk thick Pignut Hickory (Carya glabra) 5-7 leaflets Twigs smooth Buds silky Nut husk thin Pg. 11

Alternate, Compound Continued Honey Locust (Gleditsia triacanthos) Twigs, branches, and trunk have thorns up to 3 long Black Locust (Robinia pseudoacacia) Twigs, branches, and trunk have thorns up to ½ long Mimosa/Silk Tree (Albizia julibrissin) Invasive Leaves doubly compound Flowers lacy, pink in spring and summer Golden Rain-tree (Koelreuteria paniculata) Invasive 7-17 leaflets Yellow flowers in summer Seeds enclosed in lantern-like capsule Pg. 12

Opposite, Simple, Entire, Lobed Sugar Maple (Acer saccharum) 5 lobes Twigs and buds brownish Flower pale yellow Wings of fruit at right angles Norway Maple (Acer platanoides) Invasive Lobes broad Twigs and buds grayish-brown Flowers and fruit green Milky sap when broken Opposite, Simple, Entire, Not Lobed Princess Tree (Paulownia tomentosa) Invasive Leaves hairy, heart-shaped Flowers pale purple, blue in spring before leaves Catalpa sp. (Catalpa sp.) Exotic Leaves heart-shaped Flowers white with yellow purple, pink markings in spring Long seed pods through winter Flowering Dogwood (Cornus florida) Leaf veins curve to follow edge of leaf Flowers with 4 white bracts Red berries in fall Pg. 13

Opposite, Simple, Toothed, Lobed Red Maple (Acer rubrum) Twigs, buds, leafstalks reddish Flowers and fruit pink/red Silver Maple (Acer saccharinium) Twigs and buds brownish Flowers green Underside silvery-white Maple-leaved Viburnum (Viburnum acerfolium) 3 lobes Leaves coarsely-toothed Japanese Maple (Acer palmatum) Invasive 5-9 lobes Leaves red to purple in fall Cherry-leaved Viburnum (Viburnum prunifolium) Leaves resemble cherry leaves Buds brown, not hairy Opposite, Simple, Toothed, Not Lobed Arrowwood Viburnum (Viburnum dentata) Leaves coarsely-toothed Pg. 14

Opposite, Compound Buckeye sp. (Aesculus sp.) /Exotic 5 leaflets arranged in a whorled pattern Horse chestnuts (Aesculus hippocastanum) have 7-9 leaflets Box Elder (Acer negundo) 3-5 leaflets Papery fruit with wings Twigs green White Ash (Fraxinus americana) 5-9 leaflets Leaf scar is U shaped Green Ash (Fraxinus pennsylvanica) 7-9 leaflets Leaf scar is semi-circular Leaflet stalks have narrow wings Pg. 15

Index American Sycamore (Plantanus occidentalis).. Pg. 7 Ash Green (Fraxinus pennsylvanica).......... Pg. 15 White (Fraxinus americana).............. Pg. 15 Basswood, American (Tilia americana)....... Pg. 6 Beech, American (Fagus grandifolia)......... Pg. 8 Birch, River (Betula nigra).................. Pg. 7 Black Cherry (Prunus serotina).............. Pg. 8 Black Gum (Nyssa sylvatica)................ Pg. 2 Black Walnut (Juglans nigra).............. Pg. 10 Box Elder (Acer negundo)..................pg. 15 Buckeye sp. (Aesculus sp.)................. Pg. 15 Catalpa sp. (Catalpa sp.).................. Pg. 13 Chestnut American (Castanea dentata)............. Pg. 8 Chinese (Castanea mollissima)............ Pg. 8 Crape Myrtle (Lagerstroemia indica)......... Pg. 9 Eastern Redbud (Cercis canadensis)......... Pg. 5 Eastern Serviceberry (Amelanchier canadensis) Pg. 9 Elm American (Ulmus americana)............. Pg. 6 Siberian (Ulmus pumila)................. Pg. 6 Flowering Dogwood (Cornus florida)....... Pg. 13 Ginkgo (Ginkgo biloba)..................... Pg. 5 Golden Rain-tree (Koelreuteria paniculata)... Pg. 12 Hawthorn sp. (Crataegus sp.)............... Pg. 9 Hickory Bitternut (Carya cordiformis)............. Pg. 11 Mockernut (Carya tomentosa)............ Pg. 11 Pignut (Carya glabra)................... Pg. 11 Shagbark (Carya ovata)................. Pg. 11 Holly, American (Ilex opaca)............... Pg. 9 Hophornbeam (Ostrya virginiana).......... Pg. 7 Hornbeam, American (Carpinus caroliniana).. Pg. 7 Japanese Flow. Crabapple (Malus floribunda) Pg. 8 Locust Black (Robinia pseudoacacia)............. Pg. 12 Honey (Gleditsia triacanthos)............. Pg. 12 Magnolia Southern (Magnolia grandiflora)........... Pg. 1 Sweetbay (Magnolia virginiana)........... Pg. 1 Umbrella (Magnolia tripetala)............. Pg. 1 Maple Japanese (Acer palmatum)............... Pg. 14 Norway (Acer platanoides).............. Pg. 13 Red (Acer rubrum)..................... Pg. 14 Silver (Acer saccharinium)............... Pg. 14 Sugar (Acer saccharum)................. Pg. 13 Mimosa (Albizia julibrissin)............... Pg. 12 Mulberry sp. (Morus sp.)............... Pgs. 6,7,9 Oak Black (Quercus velutina)................ Pg. 3 Blackjack (Quercus marilandica).......... Pg. 3 Chestnut (Quercus montana)............. Pg. 4 Northern Red (Quercus rubra).......... Pg. 3 Pin (Quercus palustris)................. Pg. 3 Post (Quercus stellata).................. Pg. 4 Scarlet (Quercus coccinea)................pg. 3 Southern Red (Quercus falcata).......... Pg. 3. White (Quercus alba).................. Pg. 4 Willow (Quercus phellos)................ Pg. 5. Pawpaw, Common (Asimina triloba)...... Pg. 1 Pear, Bradford (Pyrus calleryana)........... Pg. 2 Persimmon, Common (Diospyros virginiana) Pg. 1 Princess Tree (Paulownia tomentosa)........pg. 13 Rhododendron sp. (Rhododendron sp.)...... Pg. 2 Sassafras (Sassafras albidum)............. Pgs.2,4 Spicebush (Lindera benzoin)............... Pg. 2 Sumac Smooth (Rhus glabra).................. Pg. 10 Staghorn (Rhus typhina)............... Pg. 10 Winged (Rhus copallina)............... Pg. 10 Sweetgum (Liquidambar styraciflua)........ Pg. 7 Tree of Heaven (Ailanthus altissima)....... Pg. 10 Tulip Poplar (Liriodendron tulipfera)........ Pg. 4 Viburnum Arrowwood (Viburnum dentata)......... Pg. 14 Cherry-leaved (Viburnum acerifolium)... Pg. 14 Maple-leaved (Viburbum prunifolium)... Pg. 14 Willow sp. (Salix sp.)..................... Pg. 9 Witch-hazel (Hamamelis virginiana)......... Pg. 6

Credits and Resources Images: American Basswood and Post Oak Images: Earl Bud Reaves. Blackjack Oak Image: The original uploader was Ed Uebel at English Wikipedia [GFDL (http://www.gnu.org/copyleft/fdl.html) or CC-BY-SA-3.0 (http://creativecommons.org/licenses/by-sa/3.0/)], via Wikimedia Commons. Hophornbeam Image: Matthew Sarver. Pignut and Shagbark Hickory Images: David Mow, https://www.flickr.com/photos/90940641@n08/with/11802696515/ Scarlet Oak Images: Evelyn Fitzgerald https://www.flickr.com/people/evelynfitzgerald/ Smooth Sumac Image: Micah Hahn, http://www.bio.brandeis.edu/fieldbio/survival/pages/about.html All images other taken by Kerry Wixted and Madeline Koenig. Resources: Kundt, J.F. & Baker, R.L. (1983). Leaf Key to Common Trees of Maryland (Extension Bulletin 238). College Park, MD: Cooperative Extension Service, University of Maryland. Maryland Plant Atlas webpage. http:// www.marylandplantatlas.org/index.php Petrides, G.A. & Wehr, Janet (1998). Peterson Field Guides: Eastern Trees. New York, NY: Houghton Mifflin Company. Sibley, D.A. (2009). The Sibley Guide to Trees. New York, NY: Alfred A. Knopf. Stein, J., Binion, D. & Acciavatta, R. (2003). Field Guide to Oak Species of Eastern North America (USDA, USFS, Publication No. FHTET-2003-01). Washington, DC: U.S. Government Printing Office. Virginia Tech Dendrology webpage. http://www.dendro.cnre.vt.edu/dendrology/main.htm Wojtech, M. (2011). Bark: A Field Guide to Trees of the Northeast. Lebanon, NH: University Press of New England. Larry Hogan, Governor Mark J. Belton, DNR Secretary dnr.maryland.gov